2006-11 – PhD,  Univ. of Helsinki & VTT, Helsinki, FI
Thesis:  Discovery of novel oxidative enzymes for food engineering. Tyrosinase and sulfhydryl oxidase / Opponent Prof. Willem van Berkel, WUR University (Wageningen, NL)

1999-05 – Master Degree in Biological Sciences (Laurea in Scienze Biologiche. mark: 110/110), Insubria University, Varese, IT
Thesis: Glycine oxidase from Bacillus subtilis, a model for the study of covalent flavinylation (my contribution was acknowledged here)

2002–03  – Erasmus exchange program, Oxford Brookes University, Oxford, UK
Courses: Plant and animal biotechnology (mark 25/30), Biochemical and molecular basis of diseases (29/30), Virology (29/30), Pharmacology (29/30), Protein and enzyme technology (29/30) Population genetics and cytogenetics (29/30), Recombinant DNA technologies (30L/30), Molecular biology of the gene (29/30) and Academic Speaking and Listening skills (mark B+)